Kingdom Plantae
The term plant is used to describe organisms that contain the following characteristics:
$ ability to synthesize carbohydrates by photosynthesis
$ presence of cellulose in cell walls
$ alternation of generations in their life cycles
$ lack of mobility
Plant Diversity
$ 300 000 to 500 000 species of identified plants
$ no one typical example of a plant
$ three broad categories of plants:
1. Aquatic plants
2. Non-vascular plants
3. Vascular plants
$ This system of classification of plants is based on the evolutionary relationships and adaptations that have occurred over million of years.
$ For example, land plants have adapted 3 primary adaptations to survive in their surroundings. These 3 adaptations are:
1. The evolution of structures and systems for the transport of water.
2. Roots, stems and leaves
3. A method of reproduction that did not depend on water for the dispersal of gametes
$ the stage of a plant that produces gametes is called a gametophyte (Haploid chromosome number).
$ the stage that produces a spore is a sporophyte (diploid chromosome number).
$ the general evolutionary trend in plants is the more complex they are the more time invested in the complex diploid form.
Aquatic Plants
$ the simplest of plants live surrounded by water because water will: provide support; keep cells from drying out; provide nutrients; and allows for the dispersal of sex cells.
$ most aquatic plants belong to one of three divisions, commonly known as green, brown and red algae. (Insert contents of table 19.2)
$ most aquatic plants are multicellular, but some are unicellular, filamentous or colonial.
$ the multi-celled body of an aquatic plant is called a thallus (plural: thalli). A thallus lacks conductive tissue as well as true roots, stems, and leaves.
Plant life on land
$ plants such as mosses, liverworts and ferns represent the beginning of what eventually evolved into larger and more complex seed bearing plants such as conifers and flowering plants.
$ when comparing plants such as mosses and fir trees, we see that there are some distinct differences.
$ Mosses are smaller and closer to the ground where as fir trees are much larger. This difference can be attributed to vascular tissue that is present in the fir trees.
$ vascular tissue or conducting tissue called xylem and phloem.
$ xylem is a vascular tissue that carries water and minerals to the leaves.
$ phloem is a vascular tissue that transports food, synthesized in the leaves to the rest of the plant.
$ plants containing these structures are called vascular plants.
$ plants without these structures are called non-vascular plants.
$ Non-vascular and vascular plants have evolved along separate lines, they share some important characteristics:
$ both contain a cuticle - a noncellular layer secreted by epidermal cells, designed to protect cells from drying out;
$ a stomata - pores in the epidermis of plants, particularly in leaves. Stomata (single: stoma) permit the exchange of gases between the plant and the atmospheric air;
$ and complex reproductive structures.
Non-Vascular ALand@ Plants
$ non-vascular land plants, or bryophytes, are confined to moist habitats because they need water for sexual reproduction to occur.
$ they live close to the ground and are abundant in wetlands, rainforests and roadside ditches.
$ there are three classes of non-vascular plants: mosses, liverworts and hornworts.
$ bryophytes are generally small plants, less than 20 cm tall.
$ they have leaf-like, stem-like and root-like structures.
$ most species have rhizoids, root like filaments that anchor the plant to the substrate.
$ the thin leaf-like structures are usually one cell layer think and are located on the top of a short main stalk. This leaf is used to absorb water and minerals and to perform photosynthesis. Water moves through the plant via cell to cell diffusion.
$ moss, which can hold large amount of water, is the most common example of a bryophyte
$ sphagnum also known as peat moss is a common example.
$ reproduction in mosses shows an interesting relationship between the sporophyte and the gametophyte generations.
$ in mosses, both generations are contained on the same plant, which is unique to land plants.
$ when the moss matures the sporangium breaks open and releases the spores.
$ when moss spores germinate it grows into a filamentous structure called a protonema, which resembles filamentous green algae
$ the protonema is the gametophyte of moss and forms the >carpet= effect
$ after several months of development, the gametophyte produces:
1. male sex organs, called antheridia (singular: antheridium) which are sperm bearing structures;
2. and female sex organs, called archegonia (singular: archegonium) which are egg-bearing structures.
$ after fertilization, the zygote develops into a short-lived sporophyte plant, and is nourished as a single stalk from the female gametophyte.
Read pages 444 - 449 for homework
Vascular Plants
$ vascular plants comprise the division Tracheophyta, which include the true terrestrial plants such as ferns, herbs, shrubs, trees and flowering plants.
$ the plants in this division transport nutrients, water and minerals via a vascular system
$ the sporophyte is the generation in which the characteristic vascular tissue, organs and systems are found.
$ the gametophyte is small, dependent on the sporophyte and short-lived.
$ with the exception of ferns, vascular plants have developed reproductive structures that are free of total dependence on water.
$ vascular tissue provides a continuous internal conduction system between the roots, stems and leaves. It also provides structural support, a feature that is essential for the large size and long life exhibited by many land plants.
$ the two main kinds of vascular plants are those that produce seeds and those that do not.
Seedless Plants
$ about 300 million years ago, seedless plants were the dominant land plants and formed extensive forests and swamp-lands.
$ these early forest and swamp-lands now make up of today=s coal mines.
$ most of these are now extinct and the modern versions are much smaller than their predecessors.
$ while adapted to life on land, seedless plants are found mainly in humid habitats because their short-lived gametophytes lack vascular tissue.
$ water is also necessary for reproduction
$ seedless plants include true ferns and whisk ferns
$ ferns are the most common of all seedless plants with just over 12 000 living species identified.
$ all ferns have conducting tissue
$ the rhizomes of most ferns are underground.
$ ferns leaves are called fronds.
$ the unfolding young fronds found in the spring are called fiddleheads
$ on the lower surface of the fronds are clusters of sporangia called sori (singular: sorus) that produce thousands of dust-like spores.
$ Mature spores are released into the air
$ a germinating spore develops into a small haploid gametophyte, called a prothallus
$ Antheridia and archegonia develop on the prothallus
$ as in bryophytes, water is needed to enable the sperm to swim to the non-motile egg.
$ following fertilization, the zygote will form a new sporophyte.
Seed Plants
$ seed plants are the most widely distributed and complex group of plants on Earth today.
$ there are more than 270 000 species of seed plants
$ first seed plants appeared about 370 million years ago
$ seed plants have separate male and female gametophytes as well as roots, stems, and leaves
$ a seed contains a plant embryo or a partially developed plant
$ many seed plants are free-living, but there are some examples of saprophytic or parasitic plants
$ most seed plants are on land, but examples such as the lilies, pitcher plants and a few trees, show that the wetlands are open to plants
$ the vascular tissue of seed plants allow them to have a larger range in size
$ there are two main seed types or groups, gymnosperms and angiosperms
$ these two words are developed from Greek, where sperma means seed, gymnos means naked, and angeion means vessel.
$ gymnosperms produce unprotected, or naked seeds, seeds in cone-like structures
$ angiosperms produce seeds that are enclosed and protected inside the fruit which is formed from the flower
$ seeds enable plants to survive by resisting dessication
Gymnosperms
$ these include pines, spruces, junipers, firs and other cone-bearing plants
$ they are characterized by thin, needle-like leaves that are specialized for hot-dry summers and cold winters and moderate rainfall
$ the needles are covered by a hard waxy cuticle that helps the plant retain moisture
$ conifers have roots that extend over a wide surface area rather than penetrate deep into the soil, this enables the tree to grow in areas where soil is thin
$ a pine tree is capable of producing millions of pollen grains which are adapted to float in the wind
$ once the pollen comes in contact with the female cone of the same tree or a different tree, the pollen grains become trapped in a sticky sap secreted by the female cone
$ as the sap dries, the pollen is drawn towards the ovule
$ the pollen produces the male sex cells
$ it is not uncommon for the time between pollenation and fertilization to take longer than a year
$ after fertilization, the resulting zygote develops into a conifer embryo.
$ the embryo and the seed coat are contained within the seed
$ the seed eventually is released from the pine cone
see diagram on page 449
Angiosperms
$ there are over 250 000 species of angiosperms, more than all other plant divisions combined
$ new species are discover almost daily
$ angiosperms are the most diverse of all the plants
$ many angiosperms are successful due to insects, a relationship that has developed over millions of years.
$ In fact, insects and some angiosperms are so interdependent that one can not live without the other.
$ This is a result of co-evolution, where they have evolved together to the point that they can not survive without each other
$ flowers are attractive to insects, birds and some mammals
$ the flower may account for the greater reproductive success of angiosperms when compared to the cones of gymnosperms
$ The Flower
$ come in all shapes and sizes, color, and give off a wide range of fragrances
$ all flowers share certain common characteristics
see page 451, fig. 19.12
$ stamen- male parts of the flower; made up of the filament, anther and pollen
1. filament- the thin stalk
2. anther- where pollen grains (male gametes) are formed
$ carpel/pistil- is the female part of the flower
1. stigma- is the sticky part on which pollen grains will land and grow.
2. style- is the slender stalk by which the pollen grains reach the ovary.
3. ovary- contains the ovule which is attached to the ovary wall by a short stalk
$ in many angiosperms, highly modified leaves, called carpels, join together to form the ovary. Inside the ovule is the embryo sac that holds the egg cell. A small hole in the wall of the ovule is called the micropyle, through which sperm nuclei enter.
$ sepals- the small green leaf-like structures that surround the carpel and stamen
$ petals- are the colorful parts of the flower that attract pollinating animals.
$ nectary- is at the base of each petal, and provides a sugary liquid called nectar
$ flowers that have carpels, stamens, petals, and sepals, such as lilies and tomatoes, are called complete flowers.
$ if one or more of these structures are lacking then they are called incomplete flowers
$ in most species, the petals die off after fertilization
$ the ovule becomes a seed after fertilization and it along with the remaining structures grow and change into fruit.
$ the seed contains the embryonic plant and the stored food for the embryo
read page 452 and 453 on reproduction of angiosperms.
Angiosperm Diversity
$ flowering plants are grouped into two classes:
1. Monocotyledons (monocots) and
2. Dicotyledons (dicots)
$ a cotyledon is a seed leaf that stores food for the young sporophyte and becomes the first leaf to appear as the seed germinates
$ common monocots include: grasses, water lilies, onions, orchids, wheat, corn, barley, and rye.
$ dicots include: maples, oaks, cacti, most forest trees and the majority of flowering plants
Page 454 questions 8-15